Dryopteris carthusiana

Family:Dryopteridaceae
Species:Dryopteris carthusiana (Vill.) Fuchs
Common Name:spinulose shield fern, spinulose wood fern, toothed wood fern
Habitat:Swamps, wet forests, and pine forests including pine plantations. Generally occurring in wet to wet-mesic soils but also in mesic to dry-mesic pine forests and also occasionally in mesic hardwood forests. It clearly has an affinity for pine plantations.
Associated Ecological Communities:**
Growth Habit:Fern
Duration:Perennial
Category:Vascular
Plant Notes:**
Taxonomic Notes:**
Status:Native, S5 (State Rank), G5 (Global Rank)
References:**

** Not applicable or data not available.

Classification

Division Pteridophyta - Ferns
Class Filicopsida
Sub Class Polypodiidae
Order Polypodiales
FamilyDryopteridaceae
Genus Dryopteris
Species Dryopteris carthusiana (Vill.) Fuchs - spinulose shield fern, spinulose wood fern, toothed wood fern

Citation

Citation Dryopteris carthusiana (Villars) H. P. Fuchs, Bull. Soc. Bot. France. 105: 339. 1959.
Basionym: Polypodium carthusianum Villars, Hist. Pl. Dauphiné 1: 292. 1786.
Type: **

** Not applicable or data not available.

Source

CountyYearHerbariaNotesSubmission Info
Albany1931 NYS    
Albany1932 NYS    
Albany1934 NYS    
Albany1949 NYS    
Albany1955 NYS    
Albany1930 NYS    
Albany1936 NYS    
Albany1933 NYS    
Albany  NYFA_1990    
Albany1931 BKL
BKL00028600
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Albany1949 BKL
BKL00028608
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Allegany  NYFA_1990    
Allegany1932 NYS    
Bronx1946 NYS    
Bronx  NYFA_1990    
Bronx1935 BKL
BKL00028615
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Cattaraugus  NYFA_1990    
Cayuga1916 NYS    
Cayuga  NYFA_1990    
Chautauqua1937 NYS    
Chautauqua  NYFA_1990    
Chemung  NYFA_1990    
Chemung1939 NYS    
Chenango  NYFA_1990    
Clinton1931 NYS    
Clinton1952 NYS    
Clinton  NYFA_1990    
Clinton1949 NYS    
Columbia  NYFA_1990    
Columbia1933 NYS    
Columbia1935 NYS    
Cortland  NYFA_1990    
Delaware1909 NYS    
Delaware1936 NYS    
Delaware  NYFA_1990    
Delaware1976 NYS    
Delaware1972 NYS    
Dutchess2001 INHS
Accession #: 211683
   
Dutchess  NYFA_1990    
Dutchess1884 NYS    
Erie  NYFA_1990    
Essex  NYFA_1990    
Essex1927 NYS    
Essex1964 NYS    
Essex1904 BKL
BKL00028592
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Franklin  NYFA_1990    
Fulton  NYFA_1990    
Fulton1959 NYS    
Genesee1924 NYS    
Genesee  NYFA_1990    
Greene1936 BKL
BKL00028599
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Greene1971 NYS    
Greene  NYFA_1990    
Greene1916 NYS    
Greene1934 NYS    
Greene1932 BKL
BKL00028597
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Hamilton1950 NYS    
Hamilton  NYFA_1990    
Herkimer1927 NYS    
Herkimer  NYFA_1990    
Jefferson1949 NYS    
Jefferson1932 NYS    
Jefferson  NYFA_1990    
Kings  NYFA_1990    
Lewis  NYFA_1990    
Livingston  NYFA_1990    
Madison  NYFA_1990    
Madison1923 NYS    
Madison1943 NYS    
Monroe1920 NYS    
Monroe  NYFA_1990    
Montgomery  NYFA_1990    
Nassau  NYFA_1990    
Nassau1992 OBPF
Log_num: 6787A
   
Nassau1907 NYS    
Nassau1941 OBPF
Log_num: 11241
Herbarium Name Used: Dryopteris spinulosa
   
Nassau1991 OBPF
Log_num: 6929
Herbarium Name Used: Dryopteris spinulosa
   
Nassau1936 BKL
BKL00028636
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Nassau2006 BKL
BKL00076615
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Nassau1920 BKL
BKL00028629
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Nassau1907 BKL
BKL00028639
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Nassau1920 BKL
BKL00028626
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Nassau1905 BKL
BKL00028632
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Nassau1915 BKL
BKL00028640
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Nassau1920 BKL
BKL00028627
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Nassau1911 BKL
BKL00028634
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Nassau1899 BKL
BKL00028633
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Nassau1916 BKL
BKL00028643
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
New York1996 BKL
BKL00028623
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
New York1996 BKL
BKL00028624
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
New York1996 BKL
BKL00028625
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
New York1995 BKL
BKL00028622
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
New York1996 NYS    
New York1995 NYS    
Niagara  NYFA_1990    
Oneida  NYFA_1990    
Oneida1901 NYS    
Oneida1899 NYS    
Onondaga  NYFA_1990    
Onondaga1919 NYS    
Ontario  NYFA_1990    
Orange  NYFA_1990    
Orange1992 NYS    
Orange1924 NYS    
Orange1991 NYS    
Orange1990 NYS    
Orange1992 CONN    
Orange1992 BKL
BKL00028619
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Orange1990 BKL
BKL00028618
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Orange1990 BKL
BKL00017817
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Orange1990 BKL
BKL00028617
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Orange1935 BKL
BKL00078454
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Orange1990 BKL
BKL00028620
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Orange1893 BKL
BKL00028604
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Orange1993 BKL
BKL00028607
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Orange1990 BKL
BKL00028756
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Orleans  NYFA_1990    
Oswego1970 RCFS
accession # 10841
Herbarium Name Used: Dryopteris carthusiana
  11/17/2009 - RCFS (Andy Nelson)
Oswego1997 RCFS
accession # 11725
Herbarium Name Used: Dryopteris carthusiana
  11/17/2009 - RCFS (Andy Nelson)
Oswego1998 RCFS
accession # 12953
Herbarium Name Used: Dryopteris carthusiana
  11/17/2009 - RCFS (Andy Nelson)
Oswego2000 RCFS
accession # 12980
Herbarium Name Used: Dryopteris carthusiana
  11/17/2009 - RCFS (Andy Nelson)
Oswego1932 NYS    
Oswego  NYFA_1990    
Oswego1998 NYS    
Oswego  NYS    
Otsego  NYFA_1990    
Putnam  NYFA_1990    
Putnam1990 BKL
BKL00028621
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Queens1886 BKL
BKL00028631
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Queens1902 BKL
BKL00028642
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Queens  NYFA_1990    
Rensselaer1920 NYS    
Rensselaer1933 NYS    
Rensselaer1997 NYS    
Rensselaer1931 NYS    
Rensselaer1999 NYS    
Rensselaer1932 NYS    
Rensselaer1996 NYS    
Rensselaer  NYFA_1990    
Richmond1905 NYS    
Richmond1908 NYS    
Richmond  NYFA_1990    
Richmond1892 BKL
BKL00028606
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Richmond1907 BKL
BKL00028595
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Richmond1904 BKL
BKL00028591
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Richmond1930 BKL
BKL00028587
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Richmond1905 BKL
BKL00028593
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Rockland1891 BKL
BKL00028596
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Rockland1935 NYS    
Rockland  NYFA_1990    
Rockland1950 NYS    
Rockland1936 CONN    
Rockland1936 NYS    
Rockland1990 NYS    
Saratoga1935 NYS    
Saratoga1931 NYS    
Saratoga1932 NYS    
Saratoga  NYFA_1990    
Saratoga1940 NYS    
Saratoga1953 NYS    
Schenectady1930 NYS    
Schenectady1950 NYS    
Schenectady  NYFA_1990    
Schoharie  NYFA_1990    
Schoharie1931 NYS    
Schuyler1991 NYS    
Schuyler  NYFA_1990    
Seneca  NYFA_1990    
Seneca1932 BKL
BKL00028594
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
St. Lawrence1990 ESF   Anne Johnson
St. Lawrence  NYFA_1990    
Steuben1932 NYS    
Steuben  NYFA_1990    
Steuben1932 BKL
BKL00028598
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Suffolk1920 BKL
BKL00028630
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Suffolk1920 BKL
BKL00028588
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Suffolk1961 BKL
BKL00080621
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Suffolk1994 BKL
BKL00028610
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Suffolk1920 BKL
BKL00028628
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Suffolk1935 BKL
BKL00028638
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Suffolk1911 BKL
BKL00028635
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Suffolk1905 BKL
BKL00028612
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Suffolk1933 BKL
BKL00028611
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Suffolk1918 BKL
BKL00028641
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Suffolk1932 BKL
BKL00028637
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Suffolk1992 CONN    
Suffolk1974 NYS    
Suffolk1929 NYS    
Suffolk1941 OBPF
Log_num: 11323
Herbarium Name Used: Dryopteris spinulosa
   
Suffolk1956 NYS    
Suffolk1973 OBPF
Log_num: 10040
Herbarium Name Used: Dryopteris spinulosa
   
Suffolk1974 OBPF
Log_num: 9197
Herbarium Name Used: Dryopteris spinulosa
   
Suffolk1990 OBPF
Log_num: 6832
Herbarium Name Used: Dryopteris spinulosa
   
Suffolk1915 NYS    
Suffolk1927 NYS    
Suffolk1990 NYS    
Suffolk1908 CONN    
Suffolk1932 NYS    
Suffolk1942 OBPF
Log_num: 11216
Herbarium Name Used: Dryopteris spinulosa
   
Suffolk1929 BKL
BKL00028609
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Suffolk1990 BKL
BKL00028616
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Suffolk1993 BKL
BKL00028563
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Suffolk  NYFA_1990    
Suffolk1928 NYS    
Sullivan1935 NYS    
Sullivan1950 NYS    
Sullivan  NYFA_1990    
Sullivan1936 NYS    
Sullivan2010 BKL
BKL00097323
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Tioga  NYFA_1990    
Tompkins  NYFA_1990    
Tompkins1935 NYS    
Tompkins1969 BKL
BKL00028586
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster1896 NYS    
Ulster1949 NYS    
Ulster1921 NYS    
Ulster  NYFA_1990    
Ulster1932 NYS    
Ulster1960 CONN
Herbarium Name Used: Dryopteris spinulosa
   
Ulster1903 NYS    
Ulster1954 NYS    
Ulster1935 NYS    
Ulster1918 BKL
BKL00028602
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2010 BKL
BKL00096773
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2010 BKL
BKL00097023
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2009 BKL
BKL00095291
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2010 BKL
BKL00097444
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2011 BKL
BKL00100812
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2011 BKL
BKL00101353
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2011 BKL
BKL00102318
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2011 BKL
BKL00101647
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2009 BKL
BKL00094722
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2009 BKL
BKL00094548
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2010 BKL
BKL00098003
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2010 BKL
BKL00097481
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2011 BKL
BKL00102322
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2011 BKL
BKL00100970
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2010 BKL
BKL00096397
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2010 BKL
BKL00098495
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2011 BKL
BKL00101992
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2009 BKL
BKL00094726
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2009 BKL
BKL00094403
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2009 BKL
BKL00095349
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster1880 BKL
BKL00028585
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2009 BKL
BKL00095779
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2009 BKL
BKL00092797
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2010 BKL
BKL00096325
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2010 BKL
BKL00098710
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2009 BKL
BKL00095644
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2009 BKL
BKL00095617
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2009 BKL
BKL00095249
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2009 BKL
BKL00095252
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Ulster2009 BKL
BKL00095526
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Warren1942 NYS    
Warren  NYFA_1990    
Warren1944 NYS    
Washington  NYFA_1990    
Washington1935 NYS    
Washington1932 BKL
BKL00028589
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Washington1928 NYS    
Washington1933 NYS    
Washington1932 BKL
BKL00028590
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Wayne1939 NYS    
Wayne  NYFA_1990    
Westchester  NYFA_1990    
Westchester1892 NYS    
Westchester1910 BKL
BKL00028613
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Westchester1991 BKL
BKL00086560
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Westchester1910 BKL
BKL00028614
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Westchester1895 BKL
BKL00028601
Herbarium Name Used: Dryopteris carthusiana
  2/8/2012 - BKL (Steve Glenn)
Wyoming  NYFA_1990    
Yates  NYFA_1990    

Synonyms

SynonymFull CitationBasionymTypeNotes
Dryopteris austriaca var. spinulosa Dryopteris austriaca (Jacq.) Woynar ex Schinz & Thellung var. spinulosa (O.F. Muell.) Fisch.   
Dryopteris spinulosa Dryopteris spinulosa (O.F. Muell.) WattPolypodium spinulosum O. F. Mueller  
Polypodium carthusianum Polypodium carthusianum Villars, Hist. Pl. Dauphiné 1: 292. 1786.   
Polypodium spinulosum Polypodium spinulosum O. F. Mueller   
Thelypteris spinulosa Thelypteris spinulosa (Mull.) Nieuwl.Polypodium carthusianum Villars, Hist. Pl. Dauphiné 1: 292. 1786.